Tiphia Fabricius 1775

Autor: Hanima, Raveendran K. P., Kumar, Girish, Hegde, Vishwanath D.
Rok vydání: 2022
Předmět:
ISSN: 0081-0282
DOI: 10.5281/zenodo.7301308
Popis: Key to the Indian subcontinent species of Tiphia (Modified from Allen, 1975; Krombein, 1982) 1. Females (antenna with 12 segments (Fig. 3); metasoma with 6 segments (Figs 6, 7); marginal cell of fore wing open (Fig. 10))................................................................................................ 2 - Males (antenna with 13 segments (Fig. 17); metasoma with 7 segments (Figs 19, 20); marginal cell of fore wing closed (Fig. 18))............................................................................................... 51 2. Apex of metasomal Gt 1 with deep, tightly infolded incision............... Tiphia (Jaynesia) assamensis Allen & Jaynes - Apex of metasomal Gt 1 without deep, tightly infolded incision (Fig. 6)........................................... 3 3. Gt1 with anterior transverse carina (Fig. 122).......................................... Tiphia (Punctotiphia) ... 4 - Gt 1 without anterior transverse carina (Fig. 299).............................................. Tiphia (Tiphia) ... 5 4. Hind basitarsus without longitudinal groove on inner face.................... Tiphia (Punctotiphia) coimbatorea Allen - Hind basitarsus with longitudinal groove on inner face (Fig. 107).................. Tiphia (Punctotiphia) exacta Nurse 5. Hind basitarsus without longitudinal groove on inner face..................................................... 6 - Hind basitarsus with longitudinal groove on inner face (Fig. 107).............................................. 26 6. Dorsal side of propodeum with conspicuous longitudinal carina about mid-way between areola and lateral margin (See Fig. 32 of Allen, 1975).............................................................. Tiphia (Tiphia) shillonga Allen - Dorsal side of propodeum without longitudinal carina between areola and lateral margin (Fig. 9)...................... 7 7. Tegula more than 1.5 × as long as middle width (Fig. 459)..................................................... 8 - Tegula at most only slightly longer than middle width (Fig. 9).................................................. 9 8. Larger species (11 mm) with black, brown to reddish-brown legs (usually coxae black) (Fig. 455)...................................................................................... Tiphia (Tiphia) quinquecarinata Cameron - Smaller species (6–8 mm) with bright-red legs (sometimes coxae black).................. Tiphia (Tiphia) tegulita Allen 9. Lateral side of propodeum without any parallel rugulae on upper part; smaller species (4.5– 6.0 mm)................................................................................................ Tiphia (Tiphia) s -tertia Allen - Lateral side of propodeum with parallel rugulae on upper part (Fig. 5); usually larger species (longer than 7 mm)........ 10 10. Femora of middle and hind legs entirely bright red to orange (Figs 432, 437)..................................... 11 - Femur of hind leg black or dull brown at least on outer face (Figs 2, 5).......................................... 14 11. Tibiae of middle and hind legs entirely bright red to orange (Figs 432, 437)...................................... 12 - Tibiae of middle and hind legs black (Figs 2, 5)............................................................ 13 12. Apical part of pygidium wrinkled and shagreened; hind tibia less strongly inflated (length 2.5 × width of tibia)................................................................................ Tiphia (Tiphia) kurczewskii Krombein - Apical part of pygidium smooth, glossy; hind tibia strongly inflated (Figs 432, 437) (length 2.1 × width of tibia).................................................................................... Tiphia (Tiphia) palmi Krombein 13. Marginal cell of fore wing nearly closed, with spur directed upward and inward toward costal margin; notaulices and anteromedian escarpment strongly connected; lower disc of punctate part of pygidium rough and almost rugose..................................................................................... Tiphia (Tiphia) scutensis Allen - Marginal cell of fore wing distinctly open, without spur or with one outwardly directed spur (Fig. 382); notaulices and anteromedian escarpment not connected (Fig. 379); lower disc of punctate part of pygidium smooth, polished........................................................................................ Tiphia (Tiphia) nathani Allen 14. Tibiae of middle and hind legs brownish yellow to dull red (Figs 52, 465, 486)................................... 15 - Tibiae of middle and hind legs black (Figs 102, 341, 344).................................................... 17 15. Submarginal carina of dorsal side of propodeum absent (Figs 56, 469); lateral side of pronotum with weak or vestigial transdiscal groove (Figs 57, 470)................................................................................ 16. - Submarginal carina of dorsal side of propodeum present (Fig. 490); lateral side of pronotum with well-developed transdiscal groove (Fig. 491)................................ Tiphia (Tiphia) sahyadriensis Hanima & Girish Kumar sp. nov. 16. Upper front mostly impunctate (Fig. 53); lateral side of propodeum weakly rugulate (Fig. 57); marginal cell and second cubital cell of fore wing with outwardly directed straight spur (Fig. 59)............... Tiphia (Tiphia) capillata Allen & Jaynes - Upper front almost uniformly punctate (Fig. 466); lateral side of propodeum strongly rugulate (Fig. 470); marginal cell and second cubital cell of fore wing without spur (Fig. 472)...... Tiphia (Tiphia) rajeevani Hanima & Girish Kumar sp. nov. 17. Areola of dorsal side of propodeum with five high, thin, parallel carinae, of which intermediate two shorter (Fig. 345)................................................................................ Tiphia (Tiphia) lyrata Magretti - Areola of dorsal side of propodeum with only three longitudinal carinae (Fig. 294)................................ 18 18. Gs 2 with anterior transverse carina; body length 17 mm........................... Tiphia (Tiphia) rothneyi Cameron - Gs2 without anterior transverse carina (Fig. 404); body length 12 mm or less..................................... 19 19. Dorsal side of pronotum with anterior transverse carina not complete, interrupted at middle (Fig. 294)................. 20 - Dorsal side of pronotum with anterior transverse carina complete (Fig. 399)...................................... 22 20. Mesoscutum with anteromedian escarpment and notaulices strongly connected (see Fig. 5 of Allen & Jaynes, 1930)............................................................................... Tiphia (Tiphia) katmanduae Allen - Mesoscutum with its anteromedian escarpment and notaulices not connected (Fig. 294)............................ 21 21. Impunctate part of pygidium conspicuously and broadly shagreened; lateral side of pronotum without transdiscal groove......................................................................... Tiphia (Tiphia) tuberculata Cameron - Impunctate part of pygidium not shagreened (Fig. 296); lateral side of pronotum with distinct transdiscal groove (Fig. 295)............................................................................ Tiphia (Tiphia) lawrencei Allen 22. Lateral side of pronotum with either very weak or without transdiscal groove (Fig. 400)............................ 23 - Lateral side of pronotum with distinct transdiscal groove (Fig. 5).............................................. 24 23. Lateral side of pronotum without transdiscal groove (Fig. 400); fore wing slightly infumated (Fig. 402)........................................................................................... Tiphia (Tiphia) nilgirensis Allen - Lateral side of pronotum with a very weak transdiscal groove; fore wing strongly yellowish.. Tiphia (Tiphia) hirsuta Smith 24. Dorsal side of propodeum without submarginal carina (Fig. 544); fore wing yellowish (Fig. 547); hind tibia inflated (2.2 × as long as broad); larger species (13–16 mm)........... Tiphia (Tiphia) venkataramani Hanima & Girish Kumar sp. nov. - Dorsal side of propodeum with submarginal carina (Figs 9, 79); fore wing hyaline or brownish infumated; hind tibia usually not inflated (2.4–3.0 × as long as broad); small to medium-sized species (5.5–10 mm)................................. 25 25. Wings yellowish hyaline (Fig. 82); carina of propodeal areola margined by obscure crenulations (Fig. 79); smaller species (5.5–6.0 mm).............................................................. Tiphia (Tiphia) cinchonae Allen - Wings brown infumated (Fig. 10); carina of propodeal areola margined by distinct crenulations (Fig. 9); medium-sized species (7–10 mm)............................................. Tiphia (Tiphia) bijui Hanima & Girish Kumar sp. nov. 26. Tegula at least 1.5 × as long as middle width (Fig. 533)...................................................... 27 - Tegula at most only slightly longer than middle width (Fig. 246)............................................... 30 27. Pronotum with vestigial anterior transverse carina present on humeral third (Fig. 533); mid leg castaneous except coxa (Figs 529, 532).................................................................. Tiphia (Tiphia) tegelonga Allen - Pronotum without anterior transverse carina; mid leg black except sometimes tarsi castaneous....................... 28 28. Legs and antennae entirely black; upper front with interspaces between punctures, on average, as wide as or wider than ocellus diameter.................................................................... Tiphia (Tiphia) oswini Turner - Legs not entirely black, sometimes fore tibia beneath red or fore and mid tarsi castaneous; antennal flagellum on lower side partly or completed castaneous; upper front with no interspaces between punctures, on average, as wide as ocellus diameter.................................................................................................... 29 29. Fore tibia beneath red; basal four flagellar segments beneath red................... Tiphia (Tiphia) knutsoni Krombein - Fore tibia beneath black; all flagellar segments beneath red........................ Tiphia (Tiphia) bouceki Krombein 30. Hind femur completely bright red (Fig. 245)............................................................... 31 - Hind femur not completely bright red, although in some species middle and hind legs are reddish at least on inner surface..................................................................................................... 37 31. Mid femur black........................................................... Tiphia (Tiphia) robusta Cameron - Mid femur red (Fig. 245).............................................................................. 32 32. Upper front without impunctate interspaces between punctures, on average, as wide as ocellus diameter................ 33 - Upper front with impunctate interspaces between punctures, on average, wider than ocellus diameter (Fig. 243)......... 34 33. Subtegular patch of microsetae wider than tegula.................................... Tiphia (Tiphia) davarae Allen - Subtegular patch of microsetae narrower than tegula............................. Tiphia (Tiphia) magrettii Cameron 34. Lateral side of pronotum with distinct transdiscal groove across its disc (Figs 247, 268)............................. 35 - Lateral side of pronotum without transdiscal groove across its disc, or if present, very weakly developed (Figs 310, 480).. 36 35. Mid and hind femora bright red (Fig. 263); mid and hind tibiae black; areola of dorsal side of propodeum without punctures (Fig. 267); apical half of pygidium polished and obscurely wrinkled (Fig. 270)........ Tiphia (Tiphia) khasiana Cameron - Mid and hind femora and tibia bright red (Fig. 242); areola of dorsal side of propodeum with punctures (Fig. 246); apical half of pygidium shagreened and apical margin wrinkled (Fig. 250)... Tiphia (Tiphia) kashmirensis Hanima & Girish Kumar 36. Hind basitarsus with very deep, narrow groove (Fig. 482); fore wing slightly brownish infumated (Fig. 483); metasoma broadly shagreened............................................................. Tiphia (Tiphia) rufofemorata Smith - Hind basitarsus with short, shallow groove, easily overlooked; fore wing yellowish hyaline (Fig. 312); metasoma polished, not shagreened...................................................... Tiphia (Tiphia) levipunctata Allen & Jaynes 37. Lateral side of pronotum without or weakly developed transdiscal groove across its disc (Fig. 138)................... 38 - Lateral side of pronotum with well-developed transdiscal groove or escarpment across its disc (Fig. 128)............... 42 38. Dorsal side of pronotum with complete anterior transverse carina present as series of short scallops (Fig. 137); hind basitarsus with short, shallow groove (Fig. 140)....................................... Tiphia (Tiphia) curvinerva Cameron - Dorsal side of pronotum with anterior transverse carina at most restricted to short distance at humeral angle (Fig. 424).... 39 39. Gt 1 with patch of dense micropunctures on anterior slope; preapical band of punctures of Gt 1 without escarpment on anterior border.................................................................... Tiphia (Tiphia) tibetana Turner - Gt1 without dense patch of micropunctures on anterior slope; preapical band of punctures of Gt1 with or without faint, crooked escarpment on its anterior border........................................................................ 40 40. Propodeal areola short, not more than 2 × as long as its apical width; preapical band of punctures of Gt 1 with faint escarpment on anterior border.................................................... Tiphia (Tiphia) pullivora Allen & Jaynes - Propodeal areola elongate, more than 2.5 × as long as its apical width (Fig. 389); preapical band of punctures of Gt 1 without trace of escarpment on anterior border (Fig. 393)........................................................... 41 41. Rugulae of lateral sides of propodeum coarse and widely separated; fore wing darkly infumated; mid tibia elongate and not inflated.................................................................... Tiphia (Tiphia) millealta Allen - Rugulae of lateral sides of propodeum shallow and closely spaced (Fig. 390); fore wing yellowish infumated (Fig. 392); mid tibia inflated on upper and outer surface.............................................. Tiphia (Tiphia) nepa Allen 42. Pygidium strongly rugose at apex (Fig. 426)............................................................... 43 - Pygidium on apical two-fifths or more smooth, not rugose or punctate (Fig. 36)................................... 44 43. Dorsal side of pronotum with strong, complete transverse carina anteriorly; scutum with anteromedian groove and notauli strongly connected; metanotum entirely coarse punctate............................ Tiphia (Tiphia) ordinaria Smith * - Dorsal side of pronotum without strong, complete transverse carina anteriorly (Fig. 424); scutum with anteromedian groove and notauli not connected (Fig. 424); metanotum laterally coarsely punctured and medially almost smooth without punctures (Fig. 424).................................................. Tiphia (Tiphia) novus Hanima & Girish Kumar sp. nov. 44. Lateral side of propodeum at least its upper part with fine or closely spaced rugulae (Fig. 128)....................... 45 - Lateral side of propodeum on upper part at least anteriorly with widely spaced rugulae (Fig. 33)...................... 46 45. Lateral side of pronotum with broad groove across its disc; metanotum usually with distantly placed punctures; marginal cell of fore wing with outwardly directed straight spur................................. Tiphia (Tiphia) s-secunda Allen - Lateral side of pronotum with deep and narrow groove across its disc (Fig. 128); metanotum with coarse punctures (Fig. 127); marginal cell of fore wing without outwardly directed straight spur (Fig. 130)............ Tiphia (Tiphia) consueta Smith 46. Groove on hind basitarsus shallow, short, vestigial........................ Tiphia (Tiphia) brevistigma Allen & Jaynes - Groove on hind basitarsus well developed (Fig. 94)......................................................... 47 47. Upper part of front with impunctate interspaces between punctures, on average, as wide as ocellus diameter (Fig. 90)......................................................................... Tiphia (Tiphia) clauseni Allen & Jaynes - Upper part of front without impunctate interspaces between punctures, on average, as wide as ocellus diameter.......... 48 48. Vertex with obscure row of minute punctures on median line extending forward from posterior aspect................. 49 - Vertex without obscure row of minute punctures on median line............................................... 50 49. Large species (Body length: 13.6–16.6 mm) with fore wing conspicuously flavous......... Tiphia (Tiphia) s-quarta Allen - Medium-sized species (Body length: 8.5–11 mm) with fore wing infumated.......... Tiphia (Tiphia) simlaensis Cameron 50. Hind tibia with its sensorium abruptly sunk below surrounding surface.................... Tiphia (Tiphia) s-sexta Allen - Hind tibia w
Published as part of Hanima, Raveendran K. P., Kumar, Girish & Hegde, Vishwanath D., 2022, Additions to the knowledge on the genus Tiphia Fabricius (Hymenoptera: Tiphiidae: Tiphiinae) from India with the description of ten new species, pp. 1-106 in Zootaxa 5204 (1) on pages 6-10, DOI: 10.11646/zootaxa.5204.1.1, http://zenodo.org/record/7285194
{"references":["Allen, H. W. (1975) The genus Tiphia of the Indian Subcontinent. Bulletin of United States Department of Agriculture, 1509, 1 - 96, 6 pls.","Krombein, K. V. (1982) Biosystematic studies of ceylonese wasps, IX. A monograph of the Tiphiidae (Hymenoptera: Vespoidea). Smithsonian contributions to Zoology, Washington, 374, 1 - 121. https: // doi. org / 10.5479 / si. 00810282.374","Allen, H. W. & Jaynes, H. A. (1930) Contribution to the taxonomy of Asiatic wasps of the genus Tiphia (Scoliidae). Proceedings of the United States National Museum, 76 (17), 1 - 105, pls. 1 - 4. https: // doi. org / 10.5479 / si. 00963801.76 - 2814.1"]}
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