Hypocaccus (Nessus) curtus Lackner & Seres 2018, comb. nov

Autor: Lackner, Tom����, Seres, G��bor
Rok vydání: 2018
Předmět:
DOI: 10.5281/zenodo.5062338
Popis: Hypocaccus (Nessus) curtus (Rosenhauer, 1847) comb. nov. (Figs 32, 40���44, 46���54) Saprinus curtus Rosenhauer, 1847: 26 (original description). MARSEUL (1855): 751 (redescription). Saprinus (Hypocaccus) curtus: GANGLBAUER (1899): 389 (redescription). Hypocacculus (Nessus) curtus: REICHARDT (1932): 49, 122 (keyed, redescription, incl. pl. IV, fig. 9); REICHARDT (1941): 285, 300 (keyed, redescription, incl. fig. 147C). Saprinus puncticollis K��ster, 1849: 30 (original description). MARSEUL (1855): 755 (redescription); BICKHARDT (1916): 96 (synonymy). Saprinus (Hypocaccus) puncticollis: GANGLBAUER (1899): 389 (redescription). Hypocacculus (Nessus) puncticollis: KRYZHANOVSKIJ & REICHARDT (1976): 204, 213 (keyed, redescription); VIENNA (1980): 179, 181 (keyed, redescription, incl. fig. 64b); MAZUR (1984): 89 (catalogue); MAZUR (1997): 254 (catalogue); Y��LAMOS (2002): 320 (keyed, redescription, incl. fig. 157f); MAZUR (2004): 94 (catalogue); MAZUR (2011): 209 (catalogue); LACKNER et al. (2015): 118 (catalogue). Saprinus cribellaticollis Jacquelin du Val, 1858:99 (original description). Fauvel in GOZIS (1886): 202 (as synonym of Saprinus puncticollis). MARSEUL (1862): 509 (redescription). Saprinus (Hypocaccus) cribellaticollis: SCHMIDT (1885): 312 (keyed). Saprinus sicanus Marseul, 1862: 490 (original description, incl. pl. XVII, fig. 47). BAUDI DI SELVE (1864): 233 (as synonym of Saprinus puncticollis). Saprinus kuesteri Marseul, 1862: 715 (catalogue; unecessary replacement name for S. puncticollis K��ster, 1849). Saprinus revisus Marseul, 1876: 39 (original description). BICKHARDT (1916): 97 (as synonym of Saprinus curtus). Type material examined. Saprinus curtus Rosenhauer, 1847. LECTOTYPE (present designation): ��� (Fig. 40), originally pinned with pin-hole in its right elytron, mounted on a rectangular mounting card, right antennal funicle and left mesotarsus missing, genitalia extracted and disarticulated, glued to the same mounting card as the specimen, ���curtus / Rosenh. [written] // Hungaria [written] // herbeus Mars. [written] // Ex Musaeo / Rosenhauer [black-margined, printed label] // pas synonime / d���Herbeus Mars. / Dr. Auzat 1917 [written-printed] // Hongrie / Ex-Musaeo / ROSENHAUER [printed] // Hypocacculus / (Nannolepidius) curtus / (Rosenhauer, 1847) / Dr. Auzat D��t. 1917 [printed] // Exemplaire provenant de la / collection Vauloger de Beaupr�� / Marcel (1862-1904) et inclus dans / la collection S. Risser en 2011 [black-margined, printed label] // Saprinus curtus / Rosenhauer, 1847 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]��� (ZSM). Saprinus puncticollis K��ster, 1849. LECTOTYPE (present designation): ��� (Fig. 42), glued onto a rectangular mounting card, two left and three right mesotarsomeres missing, genitalia extracted, disarticulated and glued to the same mounting card as the specimen, ���Typ! [written] // Cagliari / Dr. K��ster [written] // puncticollis / K��st. [written] // Saprinus / curtus Rosenh. [written] // Saprinus puncticollis / K��ster, 1849 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]��� (ZSM). Saprinus cribellaticollis Jacquelin du Val, 1858. LECTOTYPE (present designation): ♀ (Fig.41), glued on a rectangular mounting card, both antennal funicles broken off; legs: except for right foreleg and left foretibia, all tibiae broken off; with the following labels:tiny, green rectangular label that is glued onto much larger translucent plastic mounting card (original mounting card of J. du Val) and tiny, red, quadrate label, followed by, ��� Saprinus cribellaticollis / Jacquelin du Val, 1858 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]��� (MNHN; coll. Jacquelin du Val). Saprinus sicanus Marseul, 1862. LECTOTYPE (present designation):��� (Fig. 43), glued onto a rectangular mounting card, right antennal funicle, both protarsi, two segments of right mesotarsus, as well as both metatibiae missing, male genitalia extracted, disarticulated and glued onto the same mounting card as the specimen, with the following labels: small, square-shaped blue label, followed by, ��� Saprinus / sicanus m. / Schaum ���59 [round label, written] // 129c / Saprinus / sicanus m. / Sicile / Schm 679 [round label, written] // 47 (129c) Saprin / sicanus m60 / Sicil. [written] // MUSEUM PARIS / Coll. De Marseul / 2842-90 [printed] // TYPE [red-printed label; followed by: ��� Saprinus sicanus / Marseul, 1862 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]��� (MNHN). Saprinus revisus Marseul, 1876. LECTOTYPE (present designation): ♀ (Fig. 44), left antennal funicle, left protarsus, and left metatarsus missing, glued onto a rectangular mounting card, female genitalia extracted, glued to the same card as the specimen, ��� Saprinus / revisus / rest of label illegible [round, blue label, written] // MUSEUM PARIS / Coll. / De Marseul 1890 [light-green label, printed] // TYPE [red-printed label] // Saprinus revisus / Marseul, 1876 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]��� (MNHN). Additional material examined. ALGERIA: ANNABA: B��ne [= Annaba], 1 ♀, coll. Dr. Buysson (MNHN; coll. Th��rond); B��ne [= Annaba], 1 ���, Desbr. (MFNB). EGYPT: Egypt, no further data, 1 ♀, coll.Ancey, (MNHN; coll. Th��rond). FRANCE: BOUCHES- DU- RH��NE: Camargue, 2 ������, L. Puel lgt., Auzat coll. (MNHN; coll. Th��rond); Camargue, Vaccares, no date, 1 ���, 29.v.1937, 1 ���, J. Th��rond lgt. (MNHN; coll. Th��rond); Camargue, La Sauvage, 1.v.1928, 1 ���, L. Puel lgt. (MNHN; coll. Th��rond); St. Maries de la Mer, 18.vii.1922, 1 ♀, Dr. A. Chobaut lgt., coll.Dr.Auzat (MNHN; coll.Th��rond). ITALY: SARDINIA: Cagliari, Saline di Stato, 10.v.1989, 1 ���, 3 ♀♀, C. Meloni lgt. (1 ��� in CTLA, 3 ♀♀ in MSNG); Stagno di Molentargius, 27.iii.1979, 1 ���, C. Meloni lgt. (CPVV), 29.v.1988, 1 ���, 1 ♀, C. Meloni lgt. (MSNG); Serdiana, 8.vi.2003, 6 ������, 6 ♀♀, Fancello lgt. (MSNG); Molentargius, 31.i.1979, 1 ���, C. Meloni lgt. (MSNG); Cagliari, Campo Santa Gilla, 28.iii.1983, 2 ♀♀, C. Meloni lgt.(MSNG). SICILY: Sicily, no further data, 1���., 1 spec., Krtz. (MNHN); Sicilia, no further data, 1 ♀ (MFNB). LIBYA: TRIPOLI: Tripolis, no further data, 1 ♀ (MFNB). SPAIN: ANDALUSIA: Andalusia, no further data, 1♀ (MFNB). TUNISIA: TUNIS: Tunis, 1 spec., collector unknown, Reitter coll. (ZSM); Tint, i.���ii.1882, 1 ���, G. & L. Doria lgt. (ZIN); Carthage, vii. 1914, 1���, Novak lgt. (ZIN); Tunis, no further data, iv.[18]83, 1 ��� (MFNB); Tunis, no further data, 6 ������, 3 ♀♀ (MFNB); Tunis, ii.���iii.1882, 1���, G. & L. Doria lgt. (MFNB); Rad��s, iv.1933, 1���, M. Grossclaude (MNHN; coll. Th��rond). SOUSSE: Sebkha Kelbia lake near Sousse, 8.iv.1962, 1 ���, Cl. Besuchet lgt. (MSNG). Redescription. PEL: 1.60���2.00 mm; APW: 0.75���1.00 mm; PPW: 1.40���1.60 mm; EW: 1.50���1.75 mm; EL: 1.00���1.40 mm. Body (Fig. 40) oblong, oval, rather convex, cuticle dark-brown to black with faint to pronounced greenish hue; legs and antennal funicle light reddish-brown; antennal scape somewhat darker. Head: mandibles densely punctate dorsally; clypeus densely and coarsely punctate, almost rugose-lacunose, anterior margin slightly elevated; frontal disc with similar, if somewhat weaker punctation; occasionally this punctation is confluent and forms tiny rugae; frontal stria slightly outwardly arcuate, complete to reduced to interrupted medially, supraorbital stria well developed; eyes flattened, but visible from above. Basal third of frontal disc with irregular rounded glabrous area; occipital stria weak, but visible. Antennal scape somewhat darker than reddish antennal funicle, antennae similar to other species of the subgenus, sensory structures of the antennal club studied by DE MARZO & VIENNA (1982). Pronotum convex, lateral sides slightly narrowing anteriorly; anterior pronotal angles obtuse, marginal pronotal stria complete, its lateral portion observable in some cases from lateral view only. Entire pronotal disc covered with punctures separated by one to several times their diameter, punctation weakens medially. Scutellum very small, triangular. Elytra: elytral epipleuron impunctate, marginal epipleural stria complete, marginal elytral stria well developed, complete, continued as apical elytral stria for short distance. Humeral elytral stria well developed, present on basal elytral third; internal subhumeral stria present as a median fragment. Dorsal elytral striae 1���4 well developed, first the longest, slightly bisinuate, reaching approximately two-thirds of elytral length apically, occasionally even slightly longer, striae 2���4 shorter, reaching approximately elytral mid-length apically, while second stria may be longer than striae 3���4; fourth stria usually the shortest, formed in most cases of beads of punctures, stopping short of elytral mid-length apically. Fourth dorsal elytral stria usually not connected (connected in specimens that belong to form ��� cribellaticollis ���) with the basal end of (in)complete sutural elytral stria, which is in punctures and can be basally shortened. Elytral punctation covers approximately apical half of elytral disc, slightly surpassing elytral mid-length basally, slightly and scatteredly entering elytral intervals in some specimens; punctation rather dense, punctures separated by approximately their own diameter. Basal elytral fifth, fourth elytral interval, elytral flanks and extreme elytral apex impunctate, or with scattered microscopic punctation only. Propygidium and pygidium: propygidium covered with punctation similar to that of elytra; pygidium with much finer and sparser punctation. Prosternum: prosternal process slightly to moderately concave (observed from lateral view); carinal prosternal striae carinate, divergent on prosternal apophysis, running convergent to sub-parallel to almost approximate apically; from mid-length of prosternal process slightly divergent anteriorly, apically united under tiny loop; interspaces between carinal prosternal striae with scattered punctures. Lateral prosternal stria strongly carinate, convergent apically, united in front of united carinal prosternal striae; lateral sides of prosternal process densely punctate; prosternal foveae moderately large, deep. Mesoventrite: disc of mesoventrite approximately three times as wide as long, with scattered punctures (occasionally almost glabrous); marginal mesoventral stria complete, slightly inwardly arcuate medially; meso-metaventral stria undulate, bisinuate, in punctures, slightly distanced from meso-metaventral suture medially. Metaventrite: disc of metaventrite apart from several rows of tiny punctures situated along basal margin entirely glabrous; lateral metaventral stria almost straight, slightly bisinuate, deeply impressed, in punctures, stopping short of metacoxa; lateral disc of metaventrite depressed, with large oval deep punctures separated by less than their diameter; metepisternum with similar punctation, punctures of smaller sizes than those of lateral disc of metaventrite. First visible abdominal ventrite striate laterally, with scattered fine punctation, occasionally almost impunctate. Legs: protibia (Fig. 32) on outer margin with 8���11 short to moderately long denticles diminishing in size proximally, protibial groove deep; rest of leg characters similar to preceding species. Male genitalia: sternite VIII (Figs 46���47) narrowing apically; sternite VIII and tergite VIII fused laterally (Fig. 48). Tergite IX medio-laterally with tiny acute projection (Figs 49���50). Spiculum gastrale (Figs 51���52) similar to other congeners. Aedeagus (Figs 53���54) almost subparallel, bluntly pointed apically. Distribution. Hungary (?), France, Italy: Sardinia, Sicily, Spain, Portugal, Greece, Malta, Cyprus, Turkey, Tunisia, Algeria, Libya, Egypt. Biology. According to VIENNA (1980), who repeats TH��ROND (1975), H. (N.) curtus is found under detritus in sand near the seacoast, where it was collected from near Suaeda sp. and Statice virgata W. plant roots. Remarks. The type specimen was part of Rosenhauer���s collection, which later became partly a part of R. Oberth��r���s collection (A. Taghavian, pers. comm. 2017), currently housed in MNHN. The senior author has visited MNHN multiple times and failed to locate the type specimen(s) of this species in the collections of MNHN (including R. Oberth��r���s collection). Mr. Serge Risser (Pleucadeuc, France) recently purchased the Histeridae collection of the late Marcel Ren�� Paul de Vauloger de Beaupr�� and published its contents in two separate papers (RISSER 2013a,b). When reading RISSER��� S paper (2013a) we were intrigued by a specimen identified as Hypocacculus (Nannolepidius !) curtus originating from Hungary and from ���Musaeo Rosenhauer���. Mr. Risser was kind enough to send this specimen to one of us (T. L.). Having examined it as well as compared it to Rosenhauer���s original description we concluded that this is the long-lost type specimen of Rosenhauer���s species Saprinus curtus. This species was described based on an unspecified number of specimens and therefore we designate a lectotype to fix the species identity. Saprinus curtus has become a mystery practically since its description, which was, however, rather detailed and served the purpose well. The reason for this was probably the fact that the type specimen(s) were unavailable for comparison and perhaps also because no more specimens matching this species were ever reported from ���Hungary���. Based on the description alone, BICKHARDT (1916) correctly synonymized the H. (N.) puncticollis (K��ster, 1849) with H. (N.) curtus, which was also followed by REICHARDT (1932). M��LLER (1937), however, doubted the two species are synonymous since the apical elytral stria in H. (N.) curtus reaches only mid-length of elytral apex, while, according to M��LLER (1937) it is complete in H. (N.) puncticollis. Furthermore, M��LLER (1937) advocated using K��ster���s H. (N.) puncticollis as the valid (albeit not the earliest) name for this species and suggested, perhaps because of the incomplete description or the absence of the type material, that H. (N.) curtus was a dubious taxon. In the latest treatise on the Histeridae of the USSR (KRYZHANOVSKIJ & REICHARDT 1976), which in fact included almost the entire Palaearctic fauna, Kryzhanovskij upheld M��LLER��� S (1937) opinion, and the name Hypocaccus (Nessus) puncticollis gained priority. This was followed by MAZUR (1984, 1997, 2011) in all three editions of his world catalogue of the Histeridae as well as by the latest edition of the Palaearctic Catalogue by LACKNER et al. (2015). Having examined both type specimens as well as numerous non-type specimens we can conclude that the two species are synonymous, and the earlier described taxon (H. (N.) curtus) has the priority. Regarding external morphological variation of this species, see Remarks section of H. (N.) curtus . Saprinus puncticollis was described from a specimen found in Cagliari by K��ster himself, as well as from specimen(s) brought by Mr. Handschuh from Cartagena (Spain) (K��STER 1849). The depository of the Spanish specimens is unknown and hence we designate the male specimen from Cagliari (Sardinia) as the lectotype to fix the identity of this taxon for purpose of synonymy. Saprinus cribellaticollis was described based on unknown number of specimens.A single specimen was located in the original collection of Jacquelin du Val, deposited in MNHN, under the label ��� Saprinus cribellaticollis ���. Jacquelin du Val did not provide his specimens with any labels, but, according to the curator of Coleoptera in MNHN, A. Taghavian, he kept his types in his private collection. Therefore we presume that this specimen, which completely matches J. du Val���s description, is a syntype. The species was described based on an unknown number of specimens and therefore we designate the lectotype to fix the taxon identity for purpose of synonymy. Saprinus sicanus was described from Sicily (Italy) based on an unspecified number of specimens, therefore we designate the lectotype to fix the taxon identity for purpose of synonymy. Saprinus revisus was described from Algiers (Algeria) based on an unknown number of specimens, therefore we designate the lectotype to fix the taxon identity for purpose of synonymy. The type of S. curtus was found in mid-19 th century ���Hungary���. This vague locality could refer to anywhere in the former Hungarian monarchy, which stretched south to the Adriatic Sea. It is possible that this species will be discovered in countries of the former Yugoslavia. It is a rather rare and seldom-collected species apparently spread around the Mediterranean Sea. Its rarity and slight morphological differences regarding dorsal punctation or course of carinal prosternal striae probably account for its numerous synonymies. Hypocaccus (Nessus) controversus (G. M��ller, 1937) (Figs 45, 55���63) Hypocacculus controversus G. M��ller, 1937: 115 (original description). Hypocacculus (Nessus) controversus: KRYZHANOVSKIJ & REICHARDT (1976): 204, 212 (keyed, redescription); MAZUR (1984): 89 (catalogue); MAZUR (1997): 252 (catalogue); MAZUR (2004):93 (catalogue). Hypocaccus (Nessus) controversus: MAZUR (2011): 208 (catalogue); LACKNER et al. (2015): 117 (catalogue). Type material examined. Hypocacculus controversus. LECTOTYPE (present designation): ♀ (Fig. 45), mounted on a triangular mounting card, right metatarsus missing, ��� ♀ [written] // Banat 1909 / Herkulesbad / leg. M. Hilf / Coll. O. Leonhardt [printed] // sbsp. / controversus [written] // TYPUS [light-ochre label, printed] // scat. / Hist. 6 [yellow label, written] // Hypocacculus / (Nessus) / controversus / G. M��ller, 1937 / LECTOTYPE / des. T. Lackner 2017 [red label, written]��� (CST). PARALECTOTYPES: 1 ♀, side-mounted on a triangular mounting point, left meso- and metatarsus missing, ���Athen / Phaleron [written] // Da Scat. / 6 [yellow label, written] // Hypocacculus (Nessus) / controversus M��ll. / Det. T. Lackner 2017 [printed-written] // Hypocacculus / (Nessus) / controversus / G. M��ller, 1937 / PARALECTOTYPE / des. T. Lackner 2017 [red label, written]��� (CST). 1 ♀, ���
Published as part of Lackner, Tom���� & Seres, G��bor, 2018, Revision of the subgenus Nessus of the genus Hypocaccus from Central Europe, with description of a new species (Coleoptera: Histeridae), pp. 419-439 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 58 (2) on pages 432-438, DOI: 10.2478/aemnp-2018-0033, http://zenodo.org/record/4504916
{"references":["ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp.","MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758.","GANGLBAUER L. 1899: Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des franzosischen und italienischen Alpengebiets. III, 1. Familienreihe Staphylinoidea. 2 Theil: Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeritidae, Trichopterigydae, Hydroscaphidae, Scaphidiidae, Histeridae. Carl Gerolds Sohn, Wien, 408 pp.","REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164.","REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp.","KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls.","BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum. Fasc. 166 a, b. La Haye, 302 pp.","KRYZHANOVSKIJ O. L. & REICHARDT A. N. 1976: Zhuki nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). Fauna SSSR, Zhestokrylye, Tom V, vyp. 4. [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae) Fauna SSSR, Coleoptera, Vol. 5, part 4.]. Nauka, Leningrad, 434 pp (in Russian).","VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp.","MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54: 1 - 376.","MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus Supplement 7: 1 - 373.","YELAMOS T. 2002: Coleoptera, Histeridae. In: RAMOS M. A., TERCE- DOR J. A, BELLES-ROS X., GOSALBEZ-NOGUERA J., SIERRA A. G., MAYOL E. M., PIERA F. M., MARINO J. S. & GONZALEZ J. T. (eds): Fauna Iberica. Vol. 17. Museo Nacional de Ciencias Naturales, CSCI, Madrid, 411 pp.","MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. & SMETANA A. (eds): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp.","MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp.","LACKNER T., MAZUR S. & NEWTON A. F. 2015: Family Histeridae. Pp. 76 - 130. In: LOBL I. & LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Volume 1. Brill Publishers, Leiden, Boston, xxiii + 1 - 900.","JACQUELIN DU VAL M. 1858: Genere des Coleopteres d'Europe comprenant leur classification et famillies naturelles, la description de tous les genres, des tableaux synoptiques destines a faciliter l'etude, le catalogue de toutes les especes de nombreux dessins au trait ce caracteres. II. A. Deyrolle, Paris, pp. 1 - 168. Cat. pp. 53 - 83, 97 - 108.","GOZIS M. 1886: Les Histeridae Gallo-Rhenans. Tableaux traduits et abreges de l'allemand de Joh. Schmidt. Avec catalogue supplementaire par Albert Fauvel. Revue d'Entomologie 5: 152 - 213.","MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758.","SCHMIDT J. 1885: Bestimmungs-Tabellen der europaischen Coleopteren. XIV. Histeridae. Berliner Entomologische Zeitschrift 29: 279 - 330.","BAUDI DI SELVE F. 1864: Coleopterorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensito: de Europaeis notis quibusdam additis. Pars prima. Berliner Entomologische Zeitschrift 8: 195 - 233.","DE MARZO L. & VIENNA P. 1982: Osservazioni morfologiche e ultrastrutturali su particolari organi di senso delle clave antennali in Isteridi della subf. Saprininae e considerazioni sistematiche. Entomologica 17: 79 - 89.","RISSER S. 2013 a: Les Histeridae de la collection de Vauloger de Beaupre (premiere partie) (Coleoptera). L'Entomologiste 69: 197 - 214.","MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112."]}
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