Astragalus

Autor: Castillón, Eduardo Estrada, Quintanilla, José Ángel Villarreal, Salinas, Alfonso Delgado
Rok vydání: 2022
Předmět:
DOI: 10.5281/zenodo.6525302
Popis: Key to identify the different sections of Astragalus in Mexico. 1. Stipules always free or the lowest semi or almost fully amplexicaule, but never connate................................................................2 - Stipules, at least at the lowest nodes connate...................................................................................................................................20 2. Plants densely pubescent with short curly trichomes mixed with longer spirally twisted ones, appressed or spreading up to 2 mm long.....................................................................................................................................................................................................3 - Plant variously pubescent, but the trichomes 1.5 mm long or shorter, none spirally twisted............................................................4 3. Stems prostrate or decumbent, usually shorter than the longest leaves; flowers ascending; pods humistrate with age, soon caducous..................................................................................................................................................................... sect. Mollissimi - Stems erect, caulescent, the stems longer than longest leaves; flowers soon deflexed; pod erect, never humistrate, persistent................................................................................................................................................................................................ sect. Gigantei 4. Always annual; ovary (and pod) with 2 ovules only.............................................................................................. sect. Microlobium - Annual or perennial; ovary (and pod) with 4 or more ovules............................................................................................................5 5. Flowers red or, if purple, the plants acaulescent................................................................................................................................6 - Flowers purple, blue, white, lilac, concolorous, or mixtures of these colors, but never completely red; caulescent plants..............7 6. Calyx-tube shallowly campanulate, 2–4.5 mm long; banner 14 mm long or shorter; pubescence always dolabriform sect. Lotiflori - Calyx-tube cylindric or deeply campanulate, 6–10.5 mm long; banner 16–22 mm long; pubescence simple or dolabriform................................................................................................................................................................................................ sect. Argophylli 7. Pod inflated, bladder-like of papyrus texture, stiffly so or papery-membranous, unilocular or bilocular.........................................8 - Pod triquetrous or dorsoventrally flattened, not bladdery inflated, narrow elliptic, linear-oblong, oblong-elliptic, oblique-obovate or subterete in cross section, always bilocular.................................................................................................................................14 8. Flowers very small, banner 5.5–6.5 mm long; pod 4–8 mm long, persistent................................................................sect. Scalares - Flowers longer than 7 mm; pod much longer than 8 mm, caducous..................................................................................................9 9. Pod stipitate, the stipe 7–17 mm long....................................................................................................................... sect. Trichopodi - Pod sessile to subsessile, stipe (if present), 0.2-0.5 mm long..........................................................................................................10 10. Pod unilocular...................................................................................................................................................................................11 - Pod bilocular (at least below the apex or beak.................................................................................................................................12 11. Lowest stipules semi or almost fully amplexicaule......................................................................................................sect. Densifolii 11. Stipules all free...................................................................................................................................................................sect. Inflati 12. Banner 7–9.5 mm long; pod retuse in both extremes even when young, minutely apiculate and bilocular at apex.....................................................................................................................................................................................................................sect. Diphaci - Banner 10–16 mm long; pod never retuse in both extremes, the apex ending into a triangular unilocular beak............................13 13. Pod 12–23 mm long; pedicels persistent; ovules 10–42; calyx campanulate, teeth shorter than the tube...................... sect. Diphysi - Pod 6–8 mm long; pedicels caducous; ovules 4; calyx urceolate-suboblong to urceolate-globose, the teeth the same size as the tube................................................................................................................................................................................ sect. Rupertii 14. Peduncles dimorphic, the early ones very short, up to 2.6 cm long, with only 1–2 flowers, the later ones, up to 7-flowered; the banner 5–5.5 mm long; pod pendulous, sessile, subunilocular, 6–10 mm long......................................................sect. Quinqueflori - Peduncles homomorphic, if flowers are small, the root annual and pod larger, if the pod is small, the flowers more than 7-flowered; pod fully bilocular............................................................................................................................................................................15 15. Pod declined or deflexed, sessile, or elevated on a stipe-like gynophore, the body very strongly dorsoventrally compressed, peltiform or shield-shaped, as broad as long or not much longer................................................................................. sect. Scutanei - Pod ascending, spreading or deflexed, sessile or stipitate, sometimes shortly so, the body mostly lance-oblong, in profile, half ovate, in any case triquetrous or compressed triquetrous, or the body subterete to dorsoventrally flattened, linear-oblong, elliptic, oblong-elliptic, ovate-elliptic to wide-elliptic to wide-oblong, twice to many times longer than wide...........................................16 16. Pod dorsoventrally flattened, somewhat bulged, persistent on the receptacle, the body elliptic, oblong-elliptic to ovate-elliptic, straight, dorsoventrally compressed, one to one and a half longer than wide...........................................................sect. Sagitticarpi - Pod triquetrous and triangular in section or subterete, twice to many times longer than wide........................................................17 17. Pod stipitate......................................................................................................................................................................................18 - Pod sessile........................................................................................................................................................................................19 18. Pod pendulous................................................................................................................................................................. sect. Miselli - Pod incurved-ascending...................................................................................................................................... sect. Reventi-Arrecti 19. Pod deflexed; flowers very small, 4–9 mm long, early deflexed, usually very numerous (15–65)............................ sect. Micranthi - Pod ascending or loosely spreading; flowers either few or longer than 9 mm, ascending or loosely spreading.....................................................................................................................................................................................................................sect. Leptocarpi 20. Plants restricted to Baja California...................................................................................................................................................21 - Plants throughout Mexico excluding Baja California......................................................................................................................25 21. Petals large, banner 21–28 mm long; petals of the wings 19.5–25 mm long; keel petals 17.3–21 mm long; in Mexico, recorded only around Mexicali (BC)................................................................................................................................................... sect. Preusiani - Petals, all shorter; plants around Mexicali and other areas of Baja California................................................................................22 22. Pod oblong to elliptic, laterally compressed, unilocular, narrowed at both ends, basally ending in a stipe or the pod faces inflated only in the area where seeds are located..........................................................................................................................................23 - Pod widened or inflated resembling a bladder.................................................................................................................................24 23. Pod oblong to elliptic, laterally compressed, unilocular, narrowed at both ends, basally ending in a stipe.............. sect. Cusickiani - Pod inflated resembling a bladder, but its faces inflated only in the area where seeds are located..............................sect. Oxyphysi 24. Plant small, the stems up to 11 cm long; peduncles 0.5–1.5 cm long; petals small, banner 7.3–10 mm long, petals of the wings 7–9.3 mm long, keel petals 6.9–8.8 mm long; mountains of Sierra de Juárez and Sierra de San Pedro Mártir; conifer forest, 1700– 2475 m....................................................................................................................................................................... sect. Circumdati - Plants larger, the stems 30–70 cm long; peduncles 2.5–10 cm long; petals larger; coastal dunes............................ sect. Anemophili 25. Pubescence dolabriform, the trichomes joined at a point before the end, in the form of “v” or “t”, with equal or unequal ends.................................................................................................................................................................................................................26 - Pubescence basifixed with simple trichomes...................................................................................................................................27 26. Pod deflexed, bilocular, triquetrous, oblong to narrow–oblong; banner 6–8.2 mm long............................................ sect. Hypoleuci - Pod ascendant, unilocular, inflated resembling a bladder, oblong–elliptic; banner 8.3–11.8 mm long................................................................................................................................................................................................................................... sect. Humistrati 27. Petals all red; banner 22–32 mm long; petals of wings 18–31 mm largo; keel petals 18–27 mm long.....................sect. Sarcocarpi 27. Petals of another color, or if red, combined with another color; banner, wings, and keel of smaller size.......................................28 28. Pubescence of stems, petioles and peduncles dense, hispid to pilose, the trichomes straight extended or retrorse; petals purple; ovules 9–18; pod oblong to narrow-oblong, triquetrous, laterally compressed, glabrous.......................................... sect. Greggiani - Pubescence of stems, and peduncles hispid to pilose or nearly glabrous, but the trichomes appressed or ascending; pod either or different form, but if the same form, then pubescent or if trichomes extended or retrorse, then the flowers yellow......................29 29. Pod pendulous, stipitate, sometimes obscurely so............................................................................................................................30 - Pod either deflexed or ascending, always sessile.............................................................................................................................31 30. Body of the pod sharply triquetrous, unilocular; seleniferous, ill scented plants.......................................................... sect. Bisulcati - Body of the pod mostly obcompressed, dorsally excavated, unilocular to fully bilocular, a septum always present; not of seleniferous soils, scentless plants...................................................................................................................................................sect. Strigullosi 31. Body of the pod unilocular......................................................................................................................................... sect. Scytocarpi - Body of the pod semi or fully bilocular........................................................................................................................sect. Strigulosi
Published as part of Castillón, Eduardo Estrada, Quintanilla, José Ángel Villarreal & Salinas, Alfonso Delgado-, 2022, A new section of Astragalus (Fabaceae: Galegeae) from Mexico, pp. 79-86 in Phytotaxa 545 (1) on pages 83-84, DOI: 10.11646/phytotaxa.545.1.6, http://zenodo.org/record/6520428
Databáze: OpenAIRE
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