Cyrtodactylus arndti Ngo & Hormann & Pham & Phung & Ostrowski & Nguyen & Ziegler 2023, sp. nov

Autor: Ngo, Hanh Thi, Hormann, Helene, Pham, Cuong The, Phung, Trung My, Ostrowski, Sabrina, Nguyen, Truong Quang, Ziegler, Thomas
Rok vydání: 2023
Předmět:
DOI: 10.5281/zenodo.8066211
Popis: Cyrtodactylus arndti sp. nov. urn:lsid:zoobank.org:act: 83F60753-8E51-49D9-97E7-BD870AC772F3 Figs 6–9; Table 3 Diagnosis The new species of Cyrtodactylus is distinguished from remaining congeners of the C. irregularis species group by a combination of the following characters: SVL: 73.4–80.9 mm; dorsal pattern with 6 or 7 irregularly shaped bands; moderately broad nuchal band; original tail with irregular transverse bands; subcaudals transversely enlarged; 5–11 enlarged femoral scales; males with 0–2 pitted femoral pores, those absence in females; males with 6 precloacal pores, females with 6 pitted precloacal pores, pore-bearing scales arranged in a single ˄-shaped series; ventral scales 26–38; dorsal tubercles in 17–20 irregular longitudinal rows; precloacal groove absent; supralabials 8–13; infralabials 8–12; number of subdigital lamellae on fourth finger 15–20 and on fourth toe 17–22. Etymology We name this species in honor of our colleague, Prof. Dr. Hartmut Arndt, Institute of Zoology, University of Cologne, Germany, in recognition of his support for biodiversity research in Vietnam. As common names, we suggest Arndt’s Bent-toed Gecko (English) and Thằn lằn ngón arndt (Vietnamese). Type material (Figs 6–9) Holotype VIETNAM • ♂; Binh Dinh Province, Van Canh District, near Hiep Ha Village; 13°39.858′ N, 108°53.355′ E; 270 m a.s.l.; 13 Aug. 2016; D.T. Do and T.V. Nguyen; Field No. BD.2016.141; IEBR R.4930. Paratypes VIETNAM • 1 ♂; Binh Dinh Province, Quy Nhon District, Quy Nhon City; 13°41.718′ N, 109°10.277′ E; 140 m a.s.l; 8 Aug. 2016; D.T. Do and T.V. Nguyen; Field No. BD.2016.1; IEBR R.5219 • 2 ♂♂, 1 ♀; Binh Dinh Province, Van Canh District, near Dak Dum Village; 13°38.365′ N, 108°57.863′ E; 150 m a.s.l.; 11 Aug. 2016; D.T. Do and T.V. Nguyen; Field No. BD.2016.86, BD2016.87, BD.2016.88; IEBR R.4931 to IEBR.4933 • 1 ♀; Binh Dinh Province, Van Canh District, near Hiep Ha Village; 13°39.858′ N, 108°53.355′ E; 270 m a.s.l.; D.T. Do and T.V. Nguyen; Field No. BD.2016.86; IEBR R.5077 • 2 ♂♂; same collection data as for preceding; Field No. BD.2016.142, BD.2016.143; ZFMK 103910, ZFMK 103911. Description of holotype Adult male; snout-vent length 74.4 mm; tail regenerated, 98.15 mm in length; body slender, elongate (AG/SVL ratio 0.39); head distinct from neck, elongate, depressed (HL/SVL 0.29, HW/HL 0.67, HH/ HL 0.37); loreal region concave; snout long, blunt in dorsal profile (SE/HL 0.37), longer than diameter of orbit (OD/SE 0.61); scales on snout small, round or oval, granular, lager than scales on occiput; orbit large (OD/HL 0.22); pupils vertical; ear opening small, oval (ED/HL 0.06); rostral almost twice as wide as high with an inverse Y-shapted structure, surrounded by first supralabial, naris, nasorostral on each side, and internasal; nostril opening small and oval, surrounding by rostral, nasorostral, 2 supranasals and one postnasal; mental scale triangular, wider than high (ML/MW 0.74); two enlarged, triangular postmentals; supralabials 12/12; infralabials 11/13. Dorsal scales granular, dorsal tubercles round, keeled, conical, in 20 irregular rows at midbody; tubercles on occiput small; each tubercle surrounded by 9 or 10 granular scales; ventral scales smooth, round, midventral scales approximately 3–4 times as large as dorsal scales, slightly imbricate laterally, in 37 longitudinal rows at midbody between ventrolateral folds; precloacal groove absent; enlarged femoral scales 7 or 8 on each thigh, about twice the size of surrounding scales; enlarged precloacal scales 17, arranged in a rhombus; femoral pores absent; precloacal pores 6, arranged in ˄-shaped series. Fore and hindlimbs moderately slender (ForeaL/SVL 0.15 mm, CrusL/SVL 0.16); forelimbs dorsally covered by several slightly enlarged tubercles; dorsal surface of hindlimbs bearing well-developed tubercles; two postcloacal tubercles on each side on the hemipenal swellings; phalanges without webbing; each claw sheathed by two scales, the ventral sheath larger than the upper scale; number of subdigital lamellae on first finger 12/13, on first toe 13/13, on fourth finger 18/18, on fourth toe 22/22. Coloration in preservative Dorsal surface of head, body and limbs light-brown with some dark-brown pattern, without light bordering; occiput marbled with small, irregular dark-brown banded; rostral, mental, first three supralabials and first infralabials greyish brown, remaining infralabials light beige, some with greyish brown speckles or frames; nuchal band discontinuous, consisting of two stripes extending from the orbits to the neck, ending by a dark blotch on each side and a third blotch medially; dorsum with 7 irregular bands, the first two interrupted; dorsolateral region covered with round or elongate dark-brown blotches; tail with 6 dark-brown bands, fade ventrally, some small dark-brown spots arranged in a line along the lateral side of tail, tail tip dark-brown; dorsal surface of limbs with 6 or 7 irregular dark-brown bands; phalanges brown with beige knuckles; dorsal tubercles white or dark-brown depending on position; tubercles on dorsal surface of limbs and tail light-brown; venter greyish brown. Sexual dimorphism and variation The female (IEBR R.4933) differs from the males by the absence of hemipenal swellings. All male specimens have 6 precloacal pores but the females has 6 pitted scales only. Three males (IEBR R.4930, ZFMK 103910, ZFMK 103911) and the female lack femoral pores (IEBR R.5077, IEBR R.4933). For further morphological characters see Table 3. Comparisons The new species can be distinguished from all other member of Cyrtodactylus irregularis group from Vietnam by morphological characteristics (see Table 4). Cyrtodactylus arndti sp. nov. differs from C. badenensis by having the presence of enlarged femoral scales (5–11 vs absent in C. badenensis), the presence of precloacal pores in males (6 vs absent in C. badenensis), the presence of pitted precloacal pores in females (6 vs absent in C. badenensis), and different dorsal pattern (irregular bands vs banded in C. badenensis); differs from C. bidoupimontis by having the presence of pitted precloacal pores in females (6 vs absent in C. bidoupimontis), having moderately broad nuchal band, continuous or discontinuous band (vs well developed, widened posteriorly in C. bidoupimontis), different dorsal pattern (6 or 7 irregular transverse bands, colouration dark-brown on light-brown background vs 4–5 dark irregular transverse dorsal bands, usually with light borders), and the presence of transversely enlarged subcaudals vs absent in C. bidoupimontis; differs from C. bugiamapensis by having fewer precloacal pores in males (6 vs 7–11 in C. bugiamapensis), different dorsal color pattern (irregular, dark-brown transverse bands vs unclear transversal bands formed by irregular roundish to oblong, dark brown spots in C. bugiamapensis), moderately broad nuchal band, continuous or discontinuous bands (vs dark nuchal band, which can be medially divided, narrow, U-shape in C. bugiamapensis), and the presence of transversely enlarged subcaudals vs absent in C. bugiamapensis; differs from C. buchardi by having a lagger size (SVL of 73.4–80.9 vs 60–65 in C. buchardi), the presence of enlarged femoral scales (5–11 vs absent in C. buchardi), more subdigital lamellae on first finger (15–20 vs 14 in C. buchardi), more subdigital lamellae on first toe (17–22 vs 12 in C. buchardi) and the presence of transversely enlarged subcaudals vs absent in C. buchardi); differs from C. caovansungi by having a smaller size (SVL of 73.4–80.9 vs 90.4–94.0 mm), fewer femoral pores (0–2 vs 6 in C. caovansungi), fewer precloacal pores in males (6 vs 9 in C. caovansungi), the presence of pitted precloacal pores in females (6 vs absent in C. caovansungi), and fewer subdigital lamellae under the fourth toe (17–22 vs 23–25 in C. caovansungi); differs from C. cattienensis by having a larger size (73.4–80.9 mm vs 43.5–69.0 mm in C. cattienensis), the presence of pitted precloacal pores in females (6 vs absent in C. cattienensis), and the presence of transversely enlarged subcaudals (vs absent in C. cattienensis); differs from C. chungi by having a larger size (73.4–80.9 mm vs 66.6– 68.5 mm in C. chungi), fewer precloacal pores in males (6 vs 7 in C. chungi), fewer enlarged precloacal scales (17–23 vs 41–45 in C. chungi), and the presence of transversely enlarged subcaudals (vs absent in C. chungi); differs from C. cryptus by having fewer ventral scale rows (26–38 vs 47–50 in C. cryptus), the presence of enlarged femoral scales (5–11 vs absent in C. cryptus), fewer precloacal pores in males (6 vs 9–11 in C. cryptus), the presence of pitted precloacal pores in females (6 vs absent in C. cryptus), and the presence of transversely enlarged subcaudals (vs absent in C. cryptus); differs from C. cucdongensis by having a larger size (SVL 73.4–80.9 mm vs 55.8–65.9 mm in C. cucdongensis), fewer ventral scale rows (26–38 vs 41–44 in C. cucdongensis), more subdigital lamellae under the first toe (11–14 vs 8–11 in C. cucdongensis), and the presence of transversely enlarged subcaudals (vs absent in C. cucdongensis); differs from C. culaochamensis by having fewer ventral scale rows (26–38 vs 45– 50 in C. culaochamensis), the presence of enlarged femoral scales (5–11 vs absent in C. culaochamensis), fewer precloacal pores in males (6 vs 7–8 in C. culaochamensis), and the presence of pitted precloacal pores in females (6 vs absent in C. culaochamensis); differs from C. dati by having fewer ventral scale rows (26–38 vs 42–48 in C. dati), fewer femoral pores (0–2 vs 3–4 on each side in C. dati), different dorsal color pattern (irregular bands vs blotches in C. dati), and the presence of transversely enlarged subcaudals (vs absent in C. dati); differs from C. gialaiensis by having a larger size (73.4–80.9 mm vs 50.1–62.8mm in C. gialaiensis), the presence of enlarged femoral scales (5–11 vs absent in C. gialaiensis), fewer precloacal pores in males (6 vs 9–10 in C. gialaiensis), and more subdigital lamellae under the fourth finger (15–20 vs 14–15 in C. gialaiensis); differs from C. huynhi by having fewer ventral scale rows (26–38 vs 43–46 in C. huynhi), more enlarged femoral scales (5–11 vs 3–5 in C. huynhi), fewer precloacal pores in males (6 vs 7–9 in C. huynhi), and the presence of transversely enlarged subcaudals (vs absent in C. huynhi); differs from C. irregularis by having the presence of precloacal pores in males (1–2 vs absent in C. irregularis), different dorsal color pattern (irregular bands vs blotched in C. irregularis), and the presence of transversely enlarged subcaudals (vs absent in C. irregularis); differs from C. kingsadai by having a smaller size (SVL 73.4–80.9 mm vs 83.0–94.0 mm in C. kingsadai), fewer ventral scale rows (26–38 vs 39–46 in C. kingsadai), and fewer precloacal pores in males (6 vs 7–9 in C. kingsadai); differs from C. orlovi by having a larger size in males (SVL 73.4–80.9 mm vs 61.0– 68.2 mm in C. orlovi), the presence of pitted precloacal pores in females (6 vs absent in C. orlovi), and the presence of transversely enlarged subcaudals(vs absent in C. orlovi); differs from C.phnomchiensis by having fewer ventral scale rows (26–38 vs 45–54 in C. phnomchiensis), more precloacal pores in males (6 vs 4–5 in C. phnomchiensis), fewer AG/SVL ratio (0.34–0.43 vs 0.45–0.48 in C. phnomchiensis), and the presence of transversely enlarged subcaudals (vs absent in C. phnomchiensis); differs from C. phumyensis by having a larger size (SVL 73.4–80.9 mm vs 63.6–66.8 mm in C. phumyensis), and the presence of transversely enlarged subcaudals (vs absent in C. phumyensis); differs from C. phuocbinhensis by having a larger size (SVL 73.4–80.9 mm vs 46.0– 60.4 mm in C. phuocbinhensis), fewer ventral scale rows (26–38 vs 43–47 in C. phuocbinhensis), fewer precloacal pores in males (6 vs 7 in C. phuocbinhensis), the presence of pitted precloacal pores in females (6 vs absent in C. phuocbinhensis); differs from C. pseudoquadrivirgatus by having fewer ventral scale rows (26–38 vs 41–57 in C.pseudoquadrivirgatus), the presence of enlarged femoral scales (5–11 vs absent in C. pseudoquadrivirgatus), different dorsal color pattern (irregular bands vs blotched in C. pseudoquadrivirgatus), and the presence of transversely enlarged subcaudals (vs absent in C. pseudoquadrivirgatus); differs from C. raglai by having a smaller size (SVL 73.4–80.9 mm vs 95–111.7 mm in C. raglai), more precloacal pores in males (6 vs 5 in C. raglai) and the presence of pitted precloacal pores in females (6 vs absent in C. raglai); differs from C. sangi by having a larger size (SVL 73.4–80.9 mm vs 49.9–56.3 mm in C. sangi), more enlarged femoral scales (5–11 vs 4 in C. sangi), fewer precloacal pores in males (6 vs 7 in C. sangi), and more pitted precloacal pores in females (6 vs 4 in C. sangi); differs from C. takouensis by having more enlarged femoral scales (5–11 vs 3–5 in C. takouensis), more precloacal pores in males (6 vs 3–4), the presence of pitted precloacal pores in females (6 vs absent in C. takouensis), and the different dorsal color pattern (irregular bands vs banded in C. takouensis); differs from C. taynguyenensis by having more ventral scale rows (26–38 vs 42–49 in C. taynguyenensis), the presence of enlarged femoral scales (5–11 vs absent in C. taynguyenensis), the different dorsal color pattern (irregular bands vs blotched in C. taynguyenensis), and the presence of transversely enlarged subcaudals (vs absent in C. taynguyenensis); differs from C. yangbayensis by having the presence of pitted precloacal pores in females (6 vs absent in C. yangbayensis), more subdigital lamellae under the fourth toe (17–22 vs 15–17), and fewer subdigital lamellae on first toe (11–14 vs 18–20 in C. yangbayensis); differs from C. ziegleri by having a smaller size (SVL 73.4–80.9 mm vs 84.6–93.0 mm in C. ziegleri), the presence of transversely enlarged subcaudals (vs absent in C. ziegleri), and dark-brown transverse bands of the tail narrower than the light-brown interspaces (vs dark transverse bands wider than the light interspaces in C. ziegleri). Differs from C. chumuensis sp. nov. by having a larger size (SVL 73.4–80.9 mm vs maximum 67.5 mm in C. chumuensis), fewer ventral scale rows (26–38 vs 43–45 in C. chumuensis), and the presence of transversely enlarged subcaudals (vs absent in C. chumuensis). Distribution Cyrtodactylus arndti sp. nov. is currently known only from the Van Canh District, Binh Dinh Province, Vietnam (Fig. 1). Natural history Specimens were found at night between 19:00 and 22:00, on trees or on granite rock, along rocky streams, about 0.6–1.5 m above the ground, at elevations between 150 and 300 m a.s.l. The surrounding habitat was evergreen forest of medium and small hardwoods mixed with shrubs and vines (Fig. 9). The humidity was approximately 40–62% and the air temperature ranged from 28.9 to 33.1 ° C. Other reptiles species found at the sites included Acanthosaura coronata (Günther, 1861) Dixonius vietnamensis Das, 2004, Gekko gecko (Linnaeus, 1758), Gekko sp., Eutropis multifasciata (Kuhl, 1820), Boiga jaspidea (Duméril, Bibron & Duméril, 1854), Psammodynastes pulverulentus (Boie, 1827), and Trimeresurus sp.
Published as part of Ngo, Hanh Thi, Hormann, Helene, Pham, Cuong The, Phung, Trung My, Ostrowski, Sabrina, Nguyen, Truong Quang & Ziegler, Thomas, 2023, The discovery of two new species in the Cyrtodactylus irregularis group highlights that hidden diversity remains in the largest clade of the mega-diverse genus Cyrtodactylus, pp. 70-100 in European Journal of Taxonomy 875 (1) on pages 84-94, DOI: 10.5852/ejt.2023.875.2141, http://zenodo.org/record/8064396
{"references":["Ziegler T., Rosler H., Herrmann H. - W. & Vu T. N. 2002. Cyrtodactylus phongnhakebangensis sp. n., ein neuer Bogenfingergecko aus dem annamitischen Karstwaldmassiv, Vietnam. Herpetofauna 24 (141): 11 - 25.","Nguyen S. N., Orlov N. L. & Darevsky I. S. 2006. Descriptions of two new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from southern Vietnam. Russian Journal of Herpetology 13 (3): 215 - 226. https: // doi. org / 10.30906 / 1026 - 2296 - 2006 - 13 - 3 - 215 - 226","Nguyen S. N., Yang J-X., Le T-N. T., Nguyen L. T., Orlov N., Hoang C. V., Nguyen T. Q., Jin J-Q., Rao D-Q., Hoang T. N., Che J., Murphy R. W. & Zhang Y-P. 2014. DNA barcoding of Vietnamese bent-toed geckos (Squamata: Gekkonidae: Cyrtodactylus) and the description of a new species. Zootaxa 3784 (1): 48 - 66. https: // doi. org / 10.11646 / zootaxa. 3784.1.2","Nguyen S. N., Zhou W-W., Le T-N. T., Tran A-D. T., Jin J-Q., Vo B. D., Nguyen L. T., Nguyen T. T., Nguyen T. Q., Hoang D. D., Orlov N. L., Che J., Murphy R. W. & Zhang Y-P. 2017 a. Cytonuclear discordance, cryptic diversity, complex histories, and conservation needs in Vietnamese bent-toed geckos of the Cyrtodactylus irregularis species complex. Russian Journal of Herpetology 24 (2): 133 - 154.","Nguyen S. N., Nguyen L. T., Nguyen V. D. H. & Murphy R. W. 2017 b. Genetic diversity, phylogeny, and altitudinal distribution of geckos genus Cyrtodactylus on Hon Ba Mountain, Khanh Hoa Province, with note on the subcaudals of C. yangbayensis. Journal of Biotechnology 15 (3 A): 55 - 62.","Heidrich A., Rosler H., Vu N. T., Bohme W. & Ziegler T. 2007. Another new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha-Ke Bang National Park, central Truong Son, Vietnam. Zootaxa 1445: 35 - 48. https: // doi. org / 10.11646 / zootaxa. 1445.1.3","Orlov N. L., Nguyen T. Q., Nazarov A. R., Ananjeva B. N. & Nguyen S. N. 2007. A new species of the genus Cyrtodactylus Gray, 1827 and redescription of Cyrtodactylus paradoxus (Darevsky et Szczerbak, 1997) [Squamata: Sauria: Gekkonidae] from South Vietnam. Russian Journal of Herpetology 14: 145 - 152.","Nazarov R. A., Orlov N. L., Nguyen N. S. & Ho T. C. 2008. Taxonomy of naked-toe geckos Cyrtodactylus irregularis complex of South Vietnam and description of a new species from Chu Yang Sin Natural Park (Krong Bong District, Dac Lac Province), Vietnam. Russian Journal of Herpetology 15: 141 - 156.","Nazarov R., Poyarkov N. A., Orlov N. L., Phung T. M., Nguyen T. T., Hoang D. M. & Ziegler T. 2012. Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 3302 (1): 1 - 24. https: // doi. org / 10.11646 / zootaxa. 3302.1.1","Ngo T. V. & Bauer A. M. 2008. Descriptions of two new species of Cyrtodactylus Gray 1827 (Squamata: Gekkonidae) endemic to southern Vietnam. Zootaxa 1715: 27 - 42. https: // doi. org / 10.11646 / zootaxa. 1715.1.2","Ngo T. V., Grismer L. L. & Grismer J. L. 2010. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, Southwestern Vietnam. Zootaxa 2604 (1): 37 - 51. https: // doi. org / 10.11646 / zootaxa. 2604.1.3","Rosler H., Vu T. N., Nguyen T. Q., Ngo V. T. & Ziegler T. 2008. A new Cyrtodactylus (Squamata: Gekkonidae) from central Vietnam. Hamadryad 32: 125 - 140 -","Ngo T. V. & Chan K. O. 2010. A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, southern Vietnam. Zootaxa 2504 (1): 47 - 60. https: // doi. org / 10.11646 / zootaxa. 2504.1.4","Ngo T. V. & Chan O. K. 2011. A new krastic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from Northern Vietnam. Zootaxa 3125 (1): 51 - 63. https: // doi. org / 10.11646 / zootaxa. 1909.1.4","Ngo T. V. & Grismer L. L. 2012. A new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Tho Chu Island, southwestern Vietnam. Zootaxa 3228 (1): 48 - 60. https: // doi. org / 10.11646 / zootaxa. 3228.1.2","Ziegler T., Nazarov R., Orlov N., Nguyen T. Q., Vu T. N., Dang K. N., Dinh T. H. & Schmitz, A. 2010. A third new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha-Ke Bang National Park, Truong Son Range, Vietnam. Zootaxa 2413 (1): 20 - 36. https: // doi. org / 10.11646 / zootaxa. 2413.1.2","Ziegler T., Phung T. M., Le M. D. & Nguyen T. Q. 2013. A new Cyrtodactylus (Squamata: Gekkonidae) from Phu Yen Province, southern Vietnam. Zootaxa 3686 (4): 432 - 446. https: // doi. org / 10.11646 / zootaxa. 3686.4.2","Luu V. Q., Nguyen T. Q., Do Q. H. & Ziegler T. 2011. A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam. Zootaxa 3129: 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3129.1.3","Luu V. Q., Dung T. V., Nguyen T. Q., Le M. D. & Ziegler T. 2017. A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from Gia Lai Province, Central Highlands of Vietnam. Zootaxa 4362 (3): 385 - 404. https: // doi. org / 10.11646 / zootaxa. 4362.3.4","Phung T. M., van Schingen M., Ziegler T. & Nguyen T. Q. 2014. A third new Cyrtodactylus (Squamata: Gekkonidae) from Ba Den Mountain, Tay Ninh Province, southern Vietnam. Zootaxa 3764 (3): 347 - 363. https: // doi. org / 10.11646 / zootaxa. 3764.3.5","Le D. T., Nguyen T. Q., Le M. D. & Ziegler T. 2016. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Ninh Binh Province, Vietnam. Zootaxa 4162 (2): 268 - 282. https: // doi. org / 10.11646 / zootaxa. 4162.2.4","Ostrowski S., Le M. D., Ngo H. T., Pham C. T., Nguyen T. Q. & Ziegler T. 2020. A new species of Cyrtodactylus (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 4789 (1): 171 - 203. https: // doi. org / 10.11646 / zootaxa. 4789.1.5","Ostrowski S., Le M. D., Ngo H. T., Pham C. T., Phung T. M., Nguyen T. Q. & Ziegler T. 2021. A new Cyrtodactylus (Squamata: Gekkonidae) from Binh Thuan Province, southern Vietnam. European Journal of Taxonomy 731 (1): 47 - 70. https: // doi. org / 10.5852. ejt. 2021.731.1023","Do Q. H., Phung T. M., Ngo H. T., Le M. D., Ziegler T., Pham C. T. & Nguyen T. 2021. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Ninh Thuan Province, southern Vietnam. Zootaxa 4999 (1): 58 - 76. https: // doi. org / 10.11646 / zootaxa. 4999.1.4"]}
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