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Tanytydeus bazavii sp. nov. (Figs. 1–37) Diagnosis. Body large, length 600–720. Prodorsal shield bearing 2 pairs of setae (ve, sci), setae ve anterior to sci; dorsal folds between rows C–D and H–Ps strong, folds D–E, E–F, F–H showing intraspecific variability, being present, weakly formed or absent, setae c 1 23–29, c 2 38–43, d 1 24–30, e 1 21 –28, h 2 30–35; medial pore between f 1 absent. Aggenital plate with 6 pairs of setae and genital area with 8–10 pairs of setae and 3 pairs of genital papillae. Tarsi I with 1 solenidial pit distal to other. FIGUES 6–7. Tanytydeus beyzavii sp. nov. (Female): 6. Dorsal view, showing the most extreme variation in the number of folds (only dorso-sejugal, C–D, and H-Ps fold present); 7. Aggenital and genital region, showing variation in ag setae (one anterior pair absent) and asymmetry in genital setae (lacking two on one side). Description. FEMALE (holotype and 14 paratypes). Colour in life yellow. Idiosoma elongate, body constricted slightly at level of legs III–IV. Length of body (including gnathosoma) 680 (600–720); (excluding gnathosoma) 635 (550–665); width 280 (210–265); length of leg I 225 (215–230); leg II 160 (155–165); leg III 170 (160–175); leg IV 205 (200–205). Dorsum (Figs. 1, 6). Idiosoma with longitudinal striae, except for medial area of oblique to transverse striae between setal rows C and D, at level of leg III. Prodorsum with linear, crista-like shield and 3 pairs of setae (ve, sci, sce), trichobothrial setae sci and setae ve on margin of prodorsal shield, setae sce on soft cuticle, 2 pairs of peg-like setae (ep, epI) on dorsal palp coxa and coxal fields I respectively. Dorso-sejugal suture weakly to strongly formed. Opisthosoma with 6 pairs of setae and 2 pairs of cupules (im, ip); segment C with 3 pairs of setae c 1, c 2, and 1 pair of cupules (ia); all dorsal setae simple. Folds between setal rows C–D and H–Ps always present; folds between rows D–E, E–F, F–H showing intraspecific variation, being present, weakly formed or absent (e.g., compare Figs. 1, 6). Lengths of dorsal setae: ve 20 (18–28), sci 47 (44–57), sce 34 (24–34), c 1 29 (23–27), c 2 40 (38–43), d 1 30 (24–28), e 1 26 (21–28), f 1 25 (21–24), f 2 30 (27–34), h 1 31 (26–30), h 2 35 (30–33), ps 1 23 (24–28), ps 2 31 (28–33), ad 1 26 (24–25); distances between dorsal setae: ve–ve 23 (20–25), sci–sci 18 (18–23), sce–sce 80 (40–76), c 1 –c 1 44 (33–60), c 2 –c 2 205 (140–190), d 1 –d 1 85 (48–70), e 1 – e 1 88 (58–75), f 1 –f 1 56 (58–65), f 2 –f 2 205 (155–195), h 1 –h 1 63 (40–88), h 2 –h 2 150 (115–145), ve–sci 13 (9–13), sci–sce 38 (37–42), c 1 –c 2 74 (50–65), c 1 –d 1 133 (115–140), d 1 – e 1 77 (70–78), e 1 –f 1 67 (53–90), f 1 –f 2 85 (59–67), f 1 –h 1 100 (87–94), f 1 –h 2 120 (105–115), f 2 –h 2 76 (61–80), f 2 –h 1 94 (49–87), h 1 –h 2 45 (29–49). Venter (Figs. 2–3, 7) Ventral idiosoma with longitudinal striae. Endopodal shields absent. Length of setae 1a 34 (30–35), 1 b 23 (14–21), 1 c 26 (23–26), 1 d 23 (20–22), 2 b 19 (17–18), 2 c 30 (24–32), 2 d 25 (26–31), 3a 30 (31–34), 3 b 26 (20–24), 3 c 38 (31–35), 4 b 26 (21–27), 4 c 33 (21–27), ag 1 13 (12–15), ag 2 16 (14–20), ag 3 26 (25–28), ag 4 26 (24–27), ag 5 26 (24–27), ag 6 25 (24–27), g 1 16 (14–16), g 2 19 (16–20), g 3 15 (13–16), g 4 13 (13–14), g 5 12 (11–13), g 6 10 (10–12), g 7 17 (16–19), g 8 16 (14–16), g 9 12 (11–12), g 10 12 (10–11), ps 3 31 (29–32), ad 2 27 (26–28), ad 3 22 (20–23). Aggenital area with 5 or 6 pairs of setae (ag 1–6; some asymmetrical, Fig. 3): inner row with 1 or 2 pairs of setae (ag 1-2), outer row with 4 pairs (ag 3-6) almost in longitudinal row; genital plate bearing 8 to 10 pairs of genital setae (g 1-10); often asymmetrical, Fig. 7), 3 pairs of genital papillae (Va, Vm, Vp) and 3 pairs of minute setae (k 1-3) located anterior to each papilla; anus flanked by 3 pairs of setae (ad 1-3); paranal region with 3 pairs of setae, setae ps 1 dorsal and ps 2-3 ventrolateral; cupules ih anterolateral to setae ps 3. Gnathosoma (Figs. 4–5). Ventral infracapitulum with 2 pairs of subcapitular setae, m 17 (17–19), n 10 (10–11) and 2 pairs of adoral setae ao 1 8 (7–8), ao 2 6 (6–7) (Fig. 5). Palps 4 -segmented, palp trochanters nude, femoragenua with 2 setae, tibiae with 3 setae and tarsi with 4 simple setae and 3 eupathidia (Fig. 5). Chelicera 48 (44–48) long with dorsal seta and small moveable digit (Fig. 4). Legs (Figs. 8–12). Legs about one third of body length. Leg solenidia represented by pits. Setal formulae of leg segments as follows (solenidia in brackets, included in count): coxae 4 – 3 – 2 – 2; trochanters 0–1 – 1 –0; [basi]femora [4] 5 – 3 – 3 –[1] 2, genua 8 (1)– 5 (1)– 2–3; tibiae 9 (1)– 5 (1)– 4 (1)– 3; tarsi 16 (2)– 9 (1)– 6 – 5. Eupathidia: tibia I, d 1, (l); tarsus I, (ft), (tc), (p); tarsus II, pl ′. Tarsus II with bl′′ short, peg-like. MALE (7 paratypes). Body elongate. Length of body (including gnathosoma) 635–685; (excluding gnathosoma) 580–625; width 260–300; length of leg I 215–230; leg II 145–160; leg III 160–175, leg IV 180–205. Dorsum (Fig. 13). Dorsum similar to female. Lengths of dorsal setae: ve 18–28, sci 41–55, sce 30–39, c 1 24–37, c 2 29–46, d 1 25–31, e 1 24 –29, f 1 24–32, f 2 30–40, h 1 26–32, h 2 32–40. Distances between dorsal setae: ve–ve 25–28, sci–sci 16–23, sce–sce 53–80, c 1 –c 1 52–60, c 2 –c 2 145–215, d 1 –d 1 45–70, e 1 – e 1 63–75, f 1 –f 1 51–73, f 2 – f 2 163 –210, h 1 –h 1 45–62, h 2 –h 2 114–183, ve–sci 10–17, sci–sce 37–43, c 1 –c 2 59–79, c 1 –d 1 115–141, d 1 – e 1 60–75, e 1 –f 1 67–93, f 1 –f 2 67–95, f 1 –h 1 75–94, f 1 –h 2 102–118, f 2 –h 2 55–80, f 2 –h 1 80 –93, h 1 –h 2 44 –63. Venter (Figs. 14–16). Ventral idiosoma similar to female except for genitalia (Fig. 10). Lengths of ventral setae: 1a 27 – 30, 1 b 14–21, 1 c 20–26, 1 d 21–26, 2 b 16–19, 2 c 26–34, 2 d 28–35, 3 a 29–35, 3 b 17–24, 3 c 29–36, 4 b 21–26, 4 c 29–34, ps 1 23–30, ps 2 32–35, ps 3 31–36, ad 1 26–27, ad 2 26–32, ad 3 20–32. Aggenital area with 6 pairs of setae (ag 1–6): anterior 2 pairs of aggenital setae (ag 1-2) forked (Fig. 15) and rest of aggenital setae (ag 3-6) simple; genital plate bearing 12 pairs of forked setae (g 1-12), setae g 2 unpaired, 3 pairs of genital papillae. Internal genitalia with 10 pairs of setae, form as shown in Figure 16. Gnathosoma (Figs. 17–19). Ventral infracapitulum with 2 pairs of infracapitular etae m 43–51 and n 40–52, 2 pairs of adoral setae ao 1 22–27, ao 2 24–28 (Fig. 18). Palp similar to female (Fig. 19). Chelicera 43–57 long, similar to female (Fig. 18). Legs. Setae form and number same as female. TRITONYMPH (4 paratypes) Body elongate, 500–565 long, 175–215 wide; length of leg I 195–205; leg II 135–140; leg III 145–160; leg IV 175–185. Dorsum. Dorsum similar to female. All dorsal setae simple. Lengths of setae: ve 18–21, sci 48–55, sce 26–35, c 1 24–30, c 2 32–38, d 1 23–26, e 1 20 –23, f 1 29–38, f 2 30–40, h 1 26–32, h 2 31–41. Distances between dorsal setae: ve–ve 20–24, sci–sci 17–19, sce–sce 70–78, c 1 –c 1 51–56, c 2 –c 2 155–165, d 1 –d 1 46–54, e 1 – e 1 51–58, f 1 –f 1 41–52, f 2 – f 2 120–164, h 1 –h 1 35–44, h 2 –h 2 108–133, ve–sci 9–12, sci–sce 36–41, c 1 –c 2 51–57, c 1 –d 1 97–113, d 1 – e 1 51–60, e 1 –f 1 67–74, f 1 –f 2 46–58, f 1 –h 1 76–83, f 2 –h 2 52–64, h 1 –h 2 38 – 41. Venter (Fig. 20). Ventral surface same as in adult female except aggenital area with 4 pairs of setae (ag 1–4); genital plate bearing 5 pairs of genital setae (g 1-5), 3 pairs of genital papillae (Va, Vm, Vp) and 3 pairs of minute setae (k 1-3) located anterior to each papilla; anus flanked by 3 pairs of setae (ad 1-3); paranal region with 3 pairs of setae, setae ps 1 dorsal and ps 2-3 ventrolateral; cupules ih anterolateral to setae ps 3. Lengths of ventral setae: 1a 33 – 35, 1 b 18–20, 1 c 25–28, 1 d 15–22, 2 b 16–17, 2 c 27–29, 2 d 15–22, 3 a 30–33, 3 b 22–25, 3 c 27–36, 4 b 21–27, 4 c 26–30, ps 1 23–33, ps 2 29–33, ps 3 33–40, ad 1 20–25, ad 2 25–29, ad 3 25–29. Gnathosoma (Fig. 21). Ventral infracapitulum with 2 pairs of subcapitular setae m 13–18 and n 15–18, 2 pairs of adoral setae ao 1 6–8, ao 2 6–9 (Fig. 21). Palp similar to female. Legs (Figs. 22–25). Setae form and number same as adults. DEUTONYMPH (3 paratypes). Body elongate, 450–480 long, 175–180 wide; length of leg I 175–190; leg II 120–130; leg III 135–145, leg IV 155–165. Dorsum. Dorsal integument same as in adult female. Lengths of setae: ve 18–21, sci 49–50, sce 27–29, c 1 23–30, c 2 36–38, d 1 24–27, e 1 21 –24, f 1 24–30, f 2 25–30, h 1 26–32, h 2 33–41. Distances between dorsal setae: ve–ve 19–20, sci–sci 15–19, sce–sce 69–71, c 1 –c 1 43–51, c 2 –c 2 170–175, d 1 –d 1 46–55, e 1 – e 1 51–59, f 1 –f 1 50–52, f 2 – f 2 125–126, h 1 –h 1 33–36, h 2 –h 2 87–100, ve–sci 9–10, sci–sce 33–38, c 1 –c 2 45–55, c 1 –d 1 80–98, d 1 – e 1 51–59, e 1 –f 1 60–65, f 1 –f 2 45–46, f 1 –h 1 66–68, f 2 –h 2 43–50, h 1 –h 2 28 – 34. Venter (Fig. 26). Ventral idiosoma with longitudinal striae. Genital plate with 2 pairs of genital setae, 2 pairs of genital papillae (Va, Vm) and 2 pairs of minute setae (k 1-2) located anterior to each papilla; aggenital plate with 2 pairs of setae (ag 1-2). Lengths of ventral setae: 1a 31 – 35, 1 b 16–19, 1 c 26–26, 1 d 16–20, 2 b 12–15, 2 c 22–27, 2 d 30–32, 3 a 26–32, 3 b 25–26, 3 c 32–36, 4 b 18–20, 4 c 26–28, ps 1 20–23, ps 2 27–31, ps 3 35 – 34, ad 1 20–25, ad 2 25–26, ad 3 25 – 19. Gnathosoma (Fig. 27). Ventral infracapitulum with 2 pairs of subcapitular setae m 14–15 and n 11–12, 2 pairs of adoral setae ao 1 6–7, ao 2 6–9. Legs (Figs. 28–31). Setae form and number same as adults except: femur I [2] 5 (l 1 ′, l 1 ′′ absent); tibia I eupathidia l ′ (setae d 1, l′′ setiform). PROTONYMPH (1 paratype). Body elongate, 400 long, 175 wide; length of leg I 165; leg II 115; leg III 130, leg IV 135. Dorsum. Dorsum same as adult female. Lengths of setae: ve 16, sci 45, sce 22, c 1 22, c 2 34, d 1 23, e 1 18, f 1 24, f 2 30, h 1 23, h 2 30. Distances between dorsal setae: ve–ve 22, sci–sci 15, sce–sce 59, c 1 –c 1 46, c 2 –c 2 130, d 1 –d 1 44, e 1 – e 1 43, f 1 –f 1 38, f 2 – f 2 115, h 1 –h 1 38, h 2 –h 2 90, ve–sci 9, sci–sce 30, c 1 –c 2 42, d 1 – e 1 45, e 1 –f 1 52, f 1 –f 2 41, f 1 –h 1 51, h 1 –h 2 30. Venter (Fig. 32). Ventral idiosoma with longitudinal striae. Genital plate with 1 pair of genital papillae (Va) and associated eugenital seta (k 1); genital area with 1 pair of setae (ag). Lengths of ventral setae: 1a 27, 1 b 14, 1 c 13, 1 d 14, 2 c 27, 2 d 25, 3a 30, 3 b 18, 3 c 30, ps 1 19, ps 2 30, ps 3 30, ad 1 8, ad 2 16, ad 3 5. Gnathosoma (Fig. 33). Ventral infracapitulum with 2 pairs of subcapitular setae m 13 and n 7, 2 pairs of adoral setae ao 1 6, ao 2 4. Legs (Figs. 34–37). Setae form and number same as deutonymph except: coxae IV bare (setae 4 b, 4 c absent); femur IV bare; tibia IV with 2 setae (v ′ absent); tarsus IV with 3 setae ((tc) absent). LARVA. Unknown. Remarks. Tanytydeus bezavii sp. nov. resembles T. kakadu most closely by having six pairs of aggenital setae, setae ve anterior to sci and almost identical setation (Table 1). The new species differs from T. kakadu by having 8–10 pairs of genital setae (6–7 pairs in T. kakadu), a larger body size (550–685 in T. beyzavii versus 455 in T. kakadu) and longer dorsal setae (c 1 23–29, c 2 38–43, d 1 24–30, e 1 21 –28 in T. beyzavii versus c 1 15, c 2 24, d 1 14, e 1 15 in T. kakadu). Etymology. This species is named in honour of Mr G.R. Beyzavi, who kindly provided some mite specimens. Type material. Holotype femal |
