| Abstrakt: |
Highlights from the article: Bufonids, known as true toads, constitute a speciose clade of hyloid neobatrachian frogs numbering about 600 species (Frost, [16]) and with an almost cosmopolitan distribution (Pramuk et al., [30]). The oldest records were regarded as I R. i cf. I R. arenarum i (originally as cf. I R. pisanoi i , see below) and I R. i cf. I R. schneideri i , both coming from the early Pliocene Monte Hermoso Formation (Gasparini and Báez, [19]; Tomassini et al., [38]). Within the I R. marina i group, the morphology of I R. loba i suggests a close relationship with I R. arenarum i , the only extant species of the I R. marina i group currently inhabiting the South American Pampas. The late and early Pliocene ages of I R. loba i and the earliest certain records of I R. arenarum i , respectively (Pérez-Ben et al., [28]), are consistent with the estimated divergence of I R. arenarum i from its closest extant relatives during the earliest Pliocene around 5 Ma (Maciel et al., [26]). |
